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Date:         Mon, 30 Mar 2009 18:56:36 GMT
Reply-To:     Conchologists List <CONCH-L@LISTSERV.UGA.EDU>
Sender:       Conchologists List <CONCH-L@LISTSERV.UGA.EDU>
From:         Allen Aigen <serirach@JUNO.COM>
Subject:      Re: Was HAM just flim-flam?
Content-Type: text/plain; charset=windows-1252

I find it very interesting that one can essentially conceptually reverse the accepted evolution of molluscan groups. The only problem is that the fossil record does not support such a reversal, although it is rather scanty.

Allen Aigen SeriRach@juno.com

---------- Original Message ---------- From: Frederick W Schueler <bckcdb@ISTAR.CA> To: CONCH-L@LISTSERV.UGA.EDU Subject: Re: [CONCH-L] Was HAM just flim-flam? Date: Mon, 30 Mar 2009 10:24:19 -0400

Thomas Eichhorst wrote:

> I Goggled (Google Books) "ancestral mollusk fossil ham" and the first > entry that comes up is Lindberg, David R.; Ponder, Winston F.; & > Haszprunar, Gerhard. “The Mollusca: Relationships and Patterns from > Their First Half-Billion Years,” /Assembling the Tree of Life/, > (Cracraft, Joel & Donoghue, Michael J. – eds.), Oxford University Press, > pp. 252-278.

* I append a message from Taxacom that's an introduction to a recent discussion of a radical idea of molluscan ancestry. You can follow the whole thread by following the directions in the footer file.

fred. ------------------------------------------------------------ Bishops Mills Natural History Centre Frederick W. Schueler & Aleta Karstad RR#2 Bishops Mills, Ontario, Canada K0G 1T0 on the Smiths Falls Limestone Plain 44* 52'N 75* 42'W (613)258-3107 <bckcdb at istar.ca> http://pinicola.ca ------------------------------------------------------------ -------- Original Message -------- Subject: [Taxacom] Mollusca & Glossophora Date: Thu, 12 Mar 2009 21:13:18 -0500 From: kennethkinman@webtv.net (Kenneth Kinman) To: taxacom@mailman.nhm.ku.edu CC: kennethkinman@webtv.net

Dear All,

I should have noted that I do not advocate using the term "Lipocephala" instead of Bivalvia or Pelecypoda. However, Lankester's term Glossophora is a perfectly good clade according my proposed phylogeny of Mollusca. Some of the synapomorphies for Glossophora were listed with the Mollusca cladogram which I posted here on Taxacom back in 2002 (that list of preliminary synapomophies and the cladogram are as follows, with Glossophora beginning at step no. 2, between Lamellibranchia and the Euthyneura gastropods):

\________ Protobranchia 1\ \________ Lamellibranchia 2\ \___________ EUTHYNEURA \ \________Cephalopoda 3\ \____ STREPTONEURA (real torsion) 4\ \_______ Monoplacophora 5\ \______ Polyplacophora 6\ \_______ Aplacophora

1. Veliger larvae evolve.** Gills become lamellate. Crystalline style evolves.** 2. Simple radulae evolve. Thus abandon filter-feeding. Cleavage becomes unequal. Buccal development begins. Adductors reduced from 2 to 1 (or 0). Unskeletonized gills? Shells more "opisthobranch"-like.** Gastropodan muscle fine structure.** 3. Radulae become more complex, with increasing numbers of teeth per transverse row.** Mesentoblast (4d) formation begins to come after the 24-cell stage (usually 40-63). 4. "Segmentation" arises.** 8 pairs of pedal retractors. More than 6 pairs of dorsoventral muscle bundles (DVM). Over two pairs of ctenidia.** Radula bolster vesicles increase. 5. Single conch becomes multiple plates.** Spicules develop (7 rows**). Even more pairs of DVM (is not fission as likely as fusion!?) Musculature develops beneath mantle. Head appendages lost. Statocysts lost? Mesentolblast formation occurs after 63-cell stage. 6. Radular reduction. Plates lost. Muscular foot reduced. Numbers of ctenidia reduced. More than 16 prs. of DVM (certainly no reversal here). Gametes usually exit via pericardia (those which don't are probably basal aplacophorans).

NOTE: ** indicates that the synapomorphy is subject to reversals or other modifications down the line.

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