Here are a few more related titles that I know of. I index the papers in my
collection using only the authors' lastnames & abbreviated titles, but you
should be able to locate these in a library without too much difficulty. If
you can't figure out the full titles, let me know.

Aydin


Wodinsky
Am. Zoologist 9:997-1010, 1969
Penetration of shell & feeding on gastropods by Octopus

Bromley
Acta Geol. Hisp. 16:55, 1981
Concepts in ichnotaxonomy illustrated by small round holes in shells

Ambrose et al.
J. Zool. 214:491-503, 1988
Characterization of boreholes by Octopus dofleini in bivalve Saxidomus
giganteus

Bromley
Bull. Geol. Soc. Denmark 40:167, 1993
Predation habits of octopus past & present





-----Original Message-----
From: Henk and Zvia Mienis [mailto:[log in to unmask]]
Sent: Wednesday, August 08, 2001 10:51 AM
To: [log in to unmask]
Subject: Drill holes by Octopus


Dear ConchLers,

About two weeks ago a question has been posted on the Conch-L Net concerning
drill holes produced in molluscs (in particular Conidae) by Octopus. I
promised to return to the subject with some literature references. Here are
some recommended articles:

Altman, J.S. & Nixon, M., 1970. Use of the beaks and radula by Octopus
vulgaris in feeding. J. Zool., London, 161: 25-38.
Arnold, J.M. & Arnold, K.O., 1969. Some aspects of hole-boring predation by
Octopus vulgaris. Am. Zool., 9: 991-996.
Nixon, M., 1979. Hole-boring in shells by Octopus vulgaris Cuvier in the
Mediterranean. Malacology, 18: 431-443.
Nixon, M., 1979. Has Octopus vulgaris a second radula? J. Zool., London,
187: 291-296.
Nixon, M., 1980. The salivary papilla of Octopus as an accessary radula for
drilling shells. J. Zool., London, 190:53-57.
Nixon, M., 1980. The affects on shells of drilling by Octopus. J. Zool.,
London, 191: 75-88.
Pilson, M.E.Q. & Taylor, P.B., 1961. Hole drilling by Octopus. Science,
N.Y., 134: 1366-1368.
Wodinsky, J., 1973. Mechanism of hole boring in Octopus vulgaris. J. Gen.
Psychol., 88: 179-183.

Holes are drilled by Octopus with the help of chemical dissolution and
mechanical boring. The drilled holes are irregular in size and rather
perspective in form i.e. they taper considerably inwards. The penetration
hole is usually far too small to have been produced by means of the radula
of the Octopus. Fine dissection and SEM-photographs revealed to Marion Nixon
that the salivary papilla are provided with tiny teeth too as are the
posterior ends of the salivary gland duct. These rather mobile parts produce
in fact the small inner hole.

Henk K. Mienis
National Mollusc Collection
Dept. Evolution, Systematics & Ecology
Hebrew University of Jerusalem
IL-91904 Jerusalem, Israel
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